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The state legislation signed by Branstad was supported by a coalition of 35 Iowa groups opposed to abortion that contends public funding for family planning services indirectly subsidizes abortions.

The legislation represented a victory for Republican politicians who had repeatedly promised the party's rank-and-file during the 2016 election campaign that they would defund Planned Parenthood. Janet Petersen, D-Des Moines, expressed disappointment with the clinic closings.

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Planned Parenthood officials were asked by reporters Thursday if any legal challenges were anticipated regarding the state defunding legislation, but they were noncommittal.

"We will look at all of our options and weigh our legal strategies," de Baca said.

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Democrats disagreed, contending the legislation will cause Iowa women to have more difficulty obtaining birth control, cancer screenings and other health care.

In Texas, a loss of state funding by Planned Parenthood affiliates resulted in a significant increase in births among low-income women who lost access to birth control, according to a 2016 study published in the New England Journal of Medicine.

Jenifer Bowen, a spokeswoman for Iowa Right to Life, remarked, "The reallocation of our tax dollars, away from the abortion industry and into the hands of true health care facilities, will only empower more Iowa women."According to the Iowa Legislative Services Agency, there were 12,219 people participating in the state's existing family planning program in December 2016.

Republicans who supported defunding Planned Parenthood said women will have access to family planning services at 221 clinics statewide that will expand coverage to rural women who must now make long drives to urban areas for family planning services.

In three of the four counties with health centers closing — Burlington, Keokuk and Sioux City — Planned Parenthood served 80 percent or more of the family planning patients accessing care at a publicly funded provider in 2015.

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